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Using the same animal model, it was recently reported that beta aminoproprionitrile treatment induced a decrease in divalent and trivalent crosslinks, which was associated with a decrease of trabecular thickness and compressive energy to failure, while stiffness was not affected.32 Another rat study showed that increased homocysteinemia induced by a diet enriched in methiononine or homocysteine—which is likely to interfere with crosslink formation—for 3 months resulted in decreased bone formation, bone volume and bone strength at the distal femur.33 The impact of an increase of enzymatic crosslinks on bone strength can also be investigated using fetal bovine cortical bone, which is characterized by a low level of these molecules

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doi: 10.1016/j.mvr.2022.104342 126 SegelM.NeumannB.HillM