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In response to oxidative stress and ATP depletion, calcium influx opens the mitochondrial permeability transition pore (mPTP), which permits unrestricted passage of molecules up to 1.5 kDa across the inner mitochondrial membrane, leading to mitochondrial swelling, collapse of the membrane potential, cessation of ATP production, and, consequently, cell death [70]

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Total glutathione content (GSx = GSH + 2xGSSG) was determined in a Synergy 2 Multi-Mode Microplate Reader (BioTek Instruments − Vermont, USA) using the DTNB assay as described 46,47